2011g-412 2012e-555 2012e-704 animal animalbehavior behavior biologicalsciences biology chimpanzees conservation conservationofnaturalresources ecology ecosystem effectofhumanbeingson entomology japan japanesemacaque kra kyoto lifesciences macaca macacafascicularis mahalemountainsnationalpark nutrition pantroglodytes primates primatology research science socialbehavior socialbehaviorinanimals tanzania zoology